Zola was the top scientist of Johann Schmidt (Red Skull)'s Hydra science division in World War II. He was later captured by the S.S.R., then recruited by its successor, S.H.I.E.L.D. He used this opportunity to secretly rebuild Hydra from within S.H.I.E.L.D. When he suffered an incurable disease, his mind proved far too valuable, so his consciousness lived on in a super computer bunker.
Zola was actually born in Switzerland.
- Captain America: The First Avenger (2011)
- Captain America: The Winter Soldier (2014)
- Agent Carter
- 108. "Valediction" (2015)
- Avengers: Endgame (2019)